Acta Zoologica Academiae Scientiarum Hungaricae

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TAXONOMIC STUDIES ON THE GENUS PSEUDOHADENA ALPHERA KY. l889 S. L.) PART VI. DESCRIPTIONS OF EIGHT NEU TAXA OF PSEUDO HADENA S. L.) AND A NEW HETEROGRAPHA STAUDIN GER. l8J J SPECIES LEPIDOPTER A. NOCTUIDAE)'

INTRODUCTION

of important recent materiai and on the revision of sonae types, we describe severat new taxa. Simultaneo us ly, we attempt to evaluate sonae phylogeneti catly important supraspecific characters of PSeudolici lena and related genera. Based on these

Resulis of the Lepidopterological Research Project of the Himalayan wil dii se Foundation and the Hungarian Natural History Muscum sor the exploration of the satina of the Deosai NationalPark and the Northeria Areas of Pakistan. No. 3.

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characters, we try to ouiline the major cladistic relationships with in this generi cgroup and to re-evaluate the taxonomicat ranking of the supra-specific taxa. The di agnostic genital characters considered are as sol lows - Male genitalia: Shape of valva, presen ce sor reduction) of cucullus and corona, presence and specialisations os digitus sincluding secondary dyssymmetri sation), si Ze, Shape and reversibili ty of haise: shape of juxta includ-ing speciali sation of the apical pati of the juxta: Shape of the vesica, orientation. shape and si Ze of diverticula, speciali sations of the sui face of the

vesica scobination, hair- like structui es), presen ce or reduction os terminalcornutus etc. The character stati in the genera, subgenera and Species groups are considered on the Table l.

parallel lineages, e . g. in Gr&Phcintlici Or PSeui hane IcI S. Str. . the complCX, scieroti Zed plate of the antrum bursae of Eremolia lenia: the heavi ly scier- oti Zed, crenulate crest of the ductus bursae os Eremolis lenia. The broad osten hori Zontalty semiiunar, denset y setose ovi positor with wide, long falcate Sth segment and the elongate, osten retron exed. large appendix bursae with part ly scieroti Zed wrin kles, ribs or Cresis are recogniZed as

se male antrum and ductus bursae and the male vesica cata be found in many di verse fornas in ali major groups of the PSeu hia lenia generic group. The osten Similair appearance of these morphological structu res cannot be considereii as apo-morphies. In male genitalia the most important plesio morphic characters are essentiat lythe fame which also characteri Ze the 'Ground planV of Hiaden inae s. l. sinci. Hade-nini s. str. Orthosiini lenini and the hul k of the former 'UimorPhinae sensu FIBIGER & HACKER l 900): de vel oped cucullus and corona: presen ce of digitus extension of the dorso-costat scieroti sation os valva): elongate. mobile harpe. notspeciali sed clavus. uni ted willi dorsat margin os sacculus: broad. shield-shaped juxta: elongate, tubular Vesica without speciali sed sui sace structu res, but with terminal cornutus. The most typicat 'ensemble of these characters caia be observedin ' Pseudolia lena V Xantho hanes and related species, described here as Orohs dena gen. n. Oroha lenia Urimo hane x BOURSIN, l 944), O. Ciardiniati V Sp. n. , O.

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cornuti: O. Bryohia lenia subg. n.) He ΓGSOVi Sp. n. Some of these derived characters may appear as reductions os typical structures, convergent ly eVol Ved in different genera and subgenera. Such convergent

Characters are

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SYSTEMATIC PART

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Diagnosis: The closest relative of the new species is PSeu hia lenia Grgyllo-xtigma VARGA et RONKAY, l99 l) the main distinctive features are as follows: the forewing ground colour of P. albi ci Stris is paler more whitisti, the dark markings are more diffuse, tess extensive in both sex es. The male genitalia of the two sibi ing taxa are very close, the new Species haSbroader, more cui Ved harpe and more scobinate distat half of the vesica than thoseos P. GrgylisStigma VARGA et RONKAY. l99 l) the subterminal diveniculum os the Vesica projecting more to ductus Haculatorius in P. albilacustris while it is pro jected towariis the basal bulb in P. GrgylisStigmia. The female genitalia of P. albiliaci StriS are similar to P. argylloStigma but thegon ApophySeS are Shorter, the Ostium bursae is more quadratic with less roundedanterior an gles, the ductus bursae is broader, membranous, without scieroti Zed ribsat the posterior en d but with stronger anterior scobination and with smali scierot